BSc Biochemistry, Imperial College, London.

MB BS, Imperial College, London.

MSc, Institute of Psychiatry, University of London.

MD, PhD Genetics, University of London.

MRCPath, Royal College of Pathologists.

FRCPath, Royal College of Pathologists.

MRCP, Royal College of Physicians.

FRCP, Royal College of Physicians.

DSc, Imperial College, London.

Fellowship, The Academy of Medical Sciences, 2010.

Fellowship, Society of Biologists, 2011.

Fellowship European Academy of Cancer Sciences, ECCO, 2015.

FRS, The Royal Society, 2017.

British Journal of Cancer, Editor: Genetics & Genomics, 2003.

International Journal of Cancer, 2013.

scientific reports, 2015.

Cancer Family Study Group, Member, , 1992-1995; 2005-2009.

Steering Committee, Member, UKCCS, 2006.

Academic Board, Member, Institute of Cancer Research, 2001.

Joint Research Committee, Member, Institute of Cancer Research-Royal Marsden Hospital NHS Trust, 2002.

Integrating knowledge of inherited genetic variation and acquired somatic alterations in cancer tissues: promising scientific opportunities, Member, NIH, Washington, 2009-2009.

Cell and Molecular Biology Board, Board Member, MRC, 2014.